Mortoniodendron ocotense Ishiki & T. Wendt
Descripción
TREE 8–25 m tall. TRUNK (Fig. 2) 30– 60 cm d.b.h., channeled at base; BARK rugose, reddish-brown, asperous, with lenticellar warts in vertical lines; SLASH of bark yellow. BRANCHLETS 2–3 mm thick, completely covered with dense stellate trichomes, these with 8–15 arms and smaller than those of the leaf blades. LEAF BLADES coriaceous, elliptic to slightly oblong, mostly 14–23 cm long by 5.0–10.5 cm wide when mature, 2.2– 3.0 times as long as wide, pellucid-punctate due to mucilage cells 49–132 mm in diam., the blade margin entire, the blade apex gradually to abruptly acute, the acumen ca. 0.6–1.0 cm long, the base obtuse to rounded, slightly oblique; VENATION (following terminology of Leaf Architecture Working Group, 1999) weakly actinodromous with a stronger pair of suprabasal to essentially basal lateral veins (these decidedly weaker than midvein) and a much weaker exterior pair of basal veins so close to the inrolled margins as to be sometimes essentially hidden; stronger suprabasal to basal pair weakly agrophic and extending ca. 1/2 the length of the blade, diverging from midvein at narrower angle than distal secondary veins; non-basal secondary veins 5–7 per side, diverging from midvein at 45u–65u, of moderate thickness, uniformly curved, weakly to (distally) fully brochidodromous; intersecondary veins absent; tertiary veins percurrent, straight, branching 1–2 times, marginal ultimate veins looped; VESTITURE of blade adaxially very sparse with stellate trichomes or nearly glabrous, abaxially with dense persistent vestiture of stellate trichomes with 8–16(18) arms 150–300 mm long, rarely with simple (unbranched) trichomes; DOMATIA 10–13 per leaf, comprising minute glabrous areas surrounded by a ring of prominent simple hairs 1000–1300 mm long in the abaxial primary/ secondary vein axils, this ring of hairs surrounded by shorter simple or rarely stellate hairs that grade into the normal abaxial lamina hairs, occasionally in the secondary/tertiary vein axils but then the hairs smaller (450–650 mm long); STOMATA anomocytic or cyclocytic. PETIOLES of mature leaves 15–20 mm long, 1.2–2.0 mm wide, slightly thickened distally, pilose with stellate trichomes. STIPULES linear, green, dark when dry, ca. 3–4 mm long, fugacious, with scattered stellate trichomes. CYMES mostly 5–8-flowered, terminal or axillary, 5.5– 10.0 cm long, indumentum like that of the branchlets; PEDUNCLE 4–15 mm long; BRACTS 1.0–1.6 cm long, fugacious, adaxially and abaxially pilose with stellate trichomes with 12 arms; PEDICELS 12–20 mm de long, 2– 3 mm thick, distally thickened, pubescent, usually not obviously articulate above base. FLOWERS nutant, 1.2–1.4(1.7) cm long, aromatic; SEPALS 5, spreading, valvate, very thick, white (yellowish when dry), lanceolate with acute apex, 12–18 mm long, 5–7 mm wide, caducous, adaxially with six lines of simple (unbranched) trichomes, abaxially pilose with stellate trichomes like the pedicels; PETALS 5, spreading, thin, white (yellowish-red when dry), 12.5–13.0 mm long, 4 mm wide, lanceolate, with acuminate apex, caducous after anthesis, glabrous on both surfaces; STAMENS ca. 83–85, forming fascicles of 8–16, slightly inflexed, outer stamens much longer than inner, filaments white, 1–3 mm long, anthers basifixed, orange-brown, 0.6–1.1 cm long, bilocular, slightly introrse, dehiscence longitudinal, thecae 0.5–0.8 cm long.
POLLEN (Fig. 3) isopolar, oblate, P51660.00 mm, E521.4360.73 mm, P/E ratio 0.75, 3–colporate, angulaperturate, exine 1.6–2.0 mm thick, semitectate, lumen 1.2160.47 mm in diameter with 1–10 verrucae. GYNOECIUM 9–11 mm long, pubescent in basal 3/4, ovary 4–5 mm long, ovoid, 5-angled, externally densely stellate-tomentulose, the locules 5, glabrous within, the style 5–6 mm long, stellate-tomentulose in basal 2/5 to 3/5, distally glabrous, the stigma linear. FRUIT a loculicidally dehiscent woody capsule, roughly globose with 4–5 sides, ca. (3.0)5.7–6.6 cm long, ca. (2.8)4.5–6.0 cm thick (before dehiscence), externally very finely and densely stellate-tomentose, greenish brown, the outer walls thick, woody, ca. (5)7 mm thick at mid-fruit; FERTILE LOCULES 4–5, glabrous within, each with 1–3 seeds. SEED red-brown, irregularly pyriform by compression, 11 mm long; TESTA smooth, the hilum conspicuous, 1.5–2.0 mm long; ARIL orange, thin, completely covering seed.A,1
1. More detailed information on the vestiture and trichome types, on domatia, and on leaf architecture and anatomy of the new species can be found in, respectively, Rendón-Carmona et al. (2005), under the name M. ocotense, and in Solis-Montero et al. (2009), and Solis-Montero et al. (2013), under ‘‘M. sp. nov.’’ Pollen is described and illustrated in Rendón-Carmona (2005) as M. ocotense.
POLLEN (Fig. 3) isopolar, oblate, P51660.00 mm, E521.4360.73 mm, P/E ratio 0.75, 3–colporate, angulaperturate, exine 1.6–2.0 mm thick, semitectate, lumen 1.2160.47 mm in diameter with 1–10 verrucae. GYNOECIUM 9–11 mm long, pubescent in basal 3/4, ovary 4–5 mm long, ovoid, 5-angled, externally densely stellate-tomentulose, the locules 5, glabrous within, the style 5–6 mm long, stellate-tomentulose in basal 2/5 to 3/5, distally glabrous, the stigma linear. FRUIT a loculicidally dehiscent woody capsule, roughly globose with 4–5 sides, ca. (3.0)5.7–6.6 cm long, ca. (2.8)4.5–6.0 cm thick (before dehiscence), externally very finely and densely stellate-tomentose, greenish brown, the outer walls thick, woody, ca. (5)7 mm thick at mid-fruit; FERTILE LOCULES 4–5, glabrous within, each with 1–3 seeds. SEED red-brown, irregularly pyriform by compression, 11 mm long; TESTA smooth, the hilum conspicuous, 1.5–2.0 mm long; ARIL orange, thin, completely covering seed.A,1
Distribución
México (país) EndémicoA
Elevación
220 – 1070 mA
Tipo de vegetación
Selva medianaA
Categoría IUCN
En peligro (EN)B
Categoría NOM-059-SEMARNAT-2010
No incluidaC