Asplenium achilleifolium (M.Martens & Galeotti) Liebm.

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Asplenium achilleifolium (M.Martens & Galeotti) Liebm.

Descripción

Roots fibrous, not proliferous; rhizomes suberect; rhizome scales brown or often black near tips, clathrate, long-attenuate, 1.5–5 X 0.8–2 mm, entire; fronds clumped, mostly 50–120 cm; stipes greenish gray, dull, (12–)20–40 cm X 2–5 mm, 1⁄3–1⁄2 of frond length, with scattered, hair-like, clathrate scales, 0.5–1 mm long, scales 2 cells wide at bases, lacking green wings; blades chartaceous to membranaceous, ovate-lanceolate, 15–80 X 8–26 cm, 2-pinnate with wings 0.5–0.8 mm wide on pinnae, proximal pinnae slightly reduced, apices pinnatifid, nonproliferous; rachises similar to stipes in color, bearing numerous linear or hairlike scales mostly 0.5–1.5 mm long, with wings 0.3 mm wide; pinnae oblong-lanceolate, 11–18 pairs, (4–)10–15 X 2–5 cm, sessile, pinnules to ca. 20 pairs per pinna, adnate at their bases and confluent with adjacent pinnules, not or only slightly narrowed, serrately incised; veins 1 per tooth, 3–6 pairs in each pinnule, tips evident adaxially; indument abaxially of scattered, appressed, tan, clavate hairs 0.1–0.2 mm; sori 2–8 pairs per pinna; indusia 1.5–2.5 X 0.5–1 mm, margins subentire to erose or sometimes ciliate with hairs 0.1 mm long; spores reniform.A

Forma de vida

TerrestreA

Forma de crecimiento

Hierba

Nutrición

Autotrófica

Ejemplar revisado

Chis (Breedlove 69110, CAS). Col (Esperón Muñoz 18299, UC). Gro (Lorea 3020, FCME). Jal (Mexia 1252, UC). Oax (McVaugh 22363, IEB; Rzedowski 19555, NY). Pue (Lyonnet 511200001, MEXU; Riba s.n., MEXU). Ver (Purpus 4350, UC).
Unverified, Doubtful, or Mistaken Reports. Mich (reported by Mickel & Beitel, 1988, but not verified).
A

Distribución

Costa Rica presentB, El Salvador presentB, Guatemala presentB, Honduras presentB, México (Country) native and not endemicB, Nicaragua presentB, South America presentB

Elevación

(250–)5002300 mA

Ecología y Hábitat

On moist slopes in wet woods.A

Tipo de vegetación

No especificado

Categoría IUCN

No incluidaC

Categoría NOM-059-SEMARNAT-2010

No incluidaD

Estatus del taxón

(A) Como definida actualmente, probablemente una entidad natural (monofilética)

Discusión taxonómica

This highly variable species is part of a complex that involves A. blepharophorum, A. eatonii, and A. solmsii. Material of A. achilleifolium from southern Mexico falls roughly into two morphological groups: 1) specimens with large rhizome scales 4–5 X 1.5–3 mm and erose indusia 0.8–1 X 2–2.5 mm; and 2) specimens with small rhizome scales 1.5–3 X 0.5–0.8 mm and subentire indusia 0.5–0.8 X 1.5–2 mm. Specimens in this second group also have somewhat more divided pinnules, with the ultimate segments acute. They vary in the direction of A. eatonii in western Mexico. Particularly problematic are specimens from Oaxaca and Chiapas, e.g., Breedlove 24715, 24917 (DS, NY) and 31323
(DS, MEXU, NY). The interrelationships of the species in this group need clarification.
Asplenium grande, cited in synonymy, has indusia that are located along the costae where veins to the pinnules diverge, instead of on the pinnules.
Asplenium achilleifolium was reported by Mickel and Beitel (1988) from Sinaloa, but specimens from this area appear to be A. eatonii, which see.
A

Bibliografía

A. Mickel, J. T. & Smith, A. R. 2004: The Pteridophytes of Mexico Vol. 88
B. Mickel, J. T. & Smith, A. R. 2004: The Pteridophytes of Mexico Vol. 88
C. IUCN 2022: The IUCN Red List of Threatened Species. Versión 2022-2
D. SEMARNAT 2019: MODIFICACIÓN del Anexo Normativo III, Lista de especies en riesgo de la Norma Oficial Mexicana NOM-059-SEMARNAT-2010: 101 pp. – https://www.dof.gob.mx/nota_detalle.php?codigo=5578808&fecha=14/11/2019#gsc.tab=0 [accessed 2023-05-04 06:16]