Asplenium venturae A.R. Sm.
Descripción
Ab A. cristato pinnulis tantum dentatis apice, paleis rhizomatum ovatis (vs. lanceatis) apice minus acuminatis, margine dentatis dentibus 0.1–0.2 mm longis, luminibus cellularum fuscatis parvioribus et occlusioribus differt. (Honoring A. Ventura A., avid plant collector in southern Mexico and especially in the state of Veracruz.).
Roots coarse, fibrous, not proliferous; rhizomes erect; rhizome scales blackish, obscurely clathrate, lumina occluded, 2–3 X 0.5–1 mm, with marginal teeth 0.1–0.2 mm long; fronds clumped, to ca. 30 cm long; stipes dull brown, 5–12 cm X 1 mm, ca. 1⁄3–2⁄3 of frond length, narrowly green-winged adaxially, with glandtipped hairs 0.2–0.3 mm long especially abaxially, glabrescent, adaxially with wings 0.1 mm wide; blades membranaceous, 2-pinnate, 15–20 X 4–8 cm, lanceolate to deltate-oblong, apices acuminate, pinnatifid, nonproliferous; rachises brownish, glabrous, with narrow wings 0.3 mm wide; pinnae lanceate, spreading, 12–16 pairs, to 4 X 1.7 cm, sessile, with up to ca. 6 pairs of pinnules per pinna, proximal acroscopic pinnule of each pinna overlapping the rachis, usually with 1 or 2 pairs of slightly reduced proximal pinnae, pinnules rhombic to oblong, tapering to cuneate bases, 4–12 mm long, toothed toward the tips, proximal pinnules slightly lobed as well as toothed; veins mostly 2–4 pairs per pinnule, tips evident adaxially; indument abaxially of appressed to spreading gland-tipped reddish brown clavate hairs 0.1–0.2 mm; sori 1–4 per pinnule, on both sides of midveins; indusia 1–2 X 0.3–0.5 mm, margins entire; spores reniform.
A
Roots coarse, fibrous, not proliferous; rhizomes erect; rhizome scales blackish, obscurely clathrate, lumina occluded, 2–3 X 0.5–1 mm, with marginal teeth 0.1–0.2 mm long; fronds clumped, to ca. 30 cm long; stipes dull brown, 5–12 cm X 1 mm, ca. 1⁄3–2⁄3 of frond length, narrowly green-winged adaxially, with glandtipped hairs 0.2–0.3 mm long especially abaxially, glabrescent, adaxially with wings 0.1 mm wide; blades membranaceous, 2-pinnate, 15–20 X 4–8 cm, lanceolate to deltate-oblong, apices acuminate, pinnatifid, nonproliferous; rachises brownish, glabrous, with narrow wings 0.3 mm wide; pinnae lanceate, spreading, 12–16 pairs, to 4 X 1.7 cm, sessile, with up to ca. 6 pairs of pinnules per pinna, proximal acroscopic pinnule of each pinna overlapping the rachis, usually with 1 or 2 pairs of slightly reduced proximal pinnae, pinnules rhombic to oblong, tapering to cuneate bases, 4–12 mm long, toothed toward the tips, proximal pinnules slightly lobed as well as toothed; veins mostly 2–4 pairs per pinnule, tips evident adaxially; indument abaxially of appressed to spreading gland-tipped reddish brown clavate hairs 0.1–0.2 mm; sori 1–4 per pinnule, on both sides of midveins; indusia 1–2 X 0.3–0.5 mm, margins entire; spores reniform.
A
Forma de vida
TerrestreA
Ejemplar revisado
Known only from the type collection. VER (Ventura A. 11176).A
Distribución
México (Country) endemicB
Elevación
1850 mA
Ecología y Hábitat
In forests and thickets.A
Categoría IUCN
No incluidaC
Categoría NOM-059-SEMARNAT-2010
No incluidaD
Discusión taxonómica
This species is most similar to A. cristatum, but differs from that in having pinnules that are merely toothed at the tips and in having rhizome scales more ovate (vs. lanceate), with a less acuminate tips and smaller, more occluded cells, also with marginal teeth 0.1–0.2 mm long. In pinnule shape and toothing, this species is more similar to the 2-pinnate form of A. flabellulatum, but the stipes are dull brown, not lustrous, and the blades are not flagelliform at the tips. The spores appear to be well formed on the type.
A somewhat similar specimen from the same locality, Ventura A. 11171 (XAL), has larger, pinnatifid, acroscopic pinnules, reduced proximal pinnae, and narrower, more attenuate blades; the sporangia appear malformed. It could be a hybrid involving A. venturae, possibly with A. sessilifolium, under which it was first identified.
A