Asplenium blepharophorum Bertol.
Descripción
Roots coarse, fibrous, not proliferous; rhizomes erect; rhizome scales brown to tan, clathrate, 5–15 X 1–2.5 mm, with a few marginal cilia; fronds clumped, (30–)45–100 cm long; stipes brownish green to gray-brown, dull, 8–35 cm X 2–5 mm, 1⁄3 of frond length, densely clothed with clathrate scales, lacking wings;
blades chartaceous, 15–65 X 6–27 cm, usually 2-pinnatebipinnatifid, ovate-lanceolate, proximal pinnae the largest or 1–2 pairs sometimes slightly reduced, apices pinnatifid or abortive, ending in terminal buds, or buds in axils of distal pinnae, rarely buds absent; rachises yellow to gray-brown, with scattered clathrate scales 0.8–1 mm long, somewhat flattened, but not alate; pinnae 15–25 pairs, 3–14 X (1–)2–4 cm, sessile, bases truncate, sometimes narrowed on proximal pinnae, costae alate, pinnules deeply pinnatifid or pinnate (at least the larger ones), with 3–4 pairs of lobes, these obtuse to acute, sometimes bifid at tips; veins evident on both sides, tips not significantly enlarged or more visible; indument abaxially of numerous clathrate scales on costae and costules, also with tan to whitish scattered, appressed, clavate hairs 0.2–0.3 mm on veins and tissue between veins; sori 2–4 pairs per pinnule; indusia whitish, thick, 2–3 X 0.8–1 mm, margins entire or suberose; spores globose.
A
blades chartaceous, 15–65 X 6–27 cm, usually 2-pinnatebipinnatifid, ovate-lanceolate, proximal pinnae the largest or 1–2 pairs sometimes slightly reduced, apices pinnatifid or abortive, ending in terminal buds, or buds in axils of distal pinnae, rarely buds absent; rachises yellow to gray-brown, with scattered clathrate scales 0.8–1 mm long, somewhat flattened, but not alate; pinnae 15–25 pairs, 3–14 X (1–)2–4 cm, sessile, bases truncate, sometimes narrowed on proximal pinnae, costae alate, pinnules deeply pinnatifid or pinnate (at least the larger ones), with 3–4 pairs of lobes, these obtuse to acute, sometimes bifid at tips; veins evident on both sides, tips not significantly enlarged or more visible; indument abaxially of numerous clathrate scales on costae and costules, also with tan to whitish scattered, appressed, clavate hairs 0.2–0.3 mm on veins and tissue between veins; sori 2–4 pairs per pinnule; indusia whitish, thick, 2–3 X 0.8–1 mm, margins entire or suberose; spores globose.
A
Forma de vida
TerrestreA
Ejemplar revisado
Chis (Little & Sharp 9880, UC, US). Col (Sanders 10633, UC). DF (Lyonnet 3250, MEXU, US). Gro (Breedlove 65180, CAS). Hgo (Sánchez Mejorada 128, US). Jal (McVaugh 10041, DS, MEXU, NY, UC, US). Méx (Lyonnet 2090, CHAPA, MO, US). Mich (Rzedowski 41965, ENCB, IEB). Mor (Lyonnet 1862, MEXU). Oax (Pringle 6072, GH, LL, MEXU, MO, NY, UC, US). Pue (Pringle 10300, MEXU, MO, NY, UC, US). Qro (Rzedowski 48346, IEB). SLP (Virlet d’Aoust 52, P–photos MO, US). Ver (Ventura 293, DS, NY).
Unverified, Doubtful, or Mistaken Reports. Nay (reported by Mickel, 1992, but not verified; possibly the specimen seen was A. eatonii).
A
Unverified, Doubtful, or Mistaken Reports. Nay (reported by Mickel, 1992, but not verified; possibly the specimen seen was A. eatonii).
A
Elevación
1150–3100 mA
Ecología y Hábitat
Moist ravines.A
Tipo de vegetación
Bosque de neblina/mesófiloA
Categoría IUCN
No incluidaC
Categoría NOM-059-SEMARNAT-2010
No incluidaD
Discusión taxonómica
The A. blepharophorum complex, which also includes A. eatonii and A. achilleifolium, is variable with regard to the presence or absence of subterminal buds. Specimens with buds (Mickel 3810b, 4129, 4692, 4916, NY, UC), here referred to A. blepharophorum, generally have blades broadest at the bases, or the proximal pair of pinnae only slightly reduced. Of the specimens without buds, some have the proximal pinnae greatly reduced; these correspond to what is here called A. eatonii, in western Mexico. Other budless specimens have blades more or less identical to A. blepharophorum, and are here referred to that species. Asplenium achilleifolium differs in having much less scaly stipes, rachises, costae, and costules. More work is necessary to determine the relationship and status of these variations. There is an indication (large globose spores) that apogamy may be operative in A. blepharophorum.A