Asplenium miradorense Liebm.
Descripción
Roots relatively coarse, fibrous, not proliferous; rhizomes suberect to erect; rhizome scales black, weakly clathrate with occluded lumina, 1.5–3 X 0.5–0.8 mm, entire or with sparse marginal cilia; fronds clumped, mostly (10–)20–50 cm long; stipes castaneous, dull, 1–10 cm X 1–2 mm, 1⁄10–1⁄5 of frond length, glabrous except for scales at bases, not winged; blades chartaceous, 1-pinnate, 25–45(–55) X 3–5(–7) cm, linear-oblong, apices pinnatifid, not proliferous; rachises castaneous, dull, glabrous, with greenish wings 0.3–0.5 mm wide; pinnae oblong-lanceolate, 16–55 pairs, reduced proximally, 1.5–3 X 0.5–1 cm, sessile or stalked to 1 mm, inequilateral, basiscopically excavate, acroscopic margins straight to curved, bases often overlapping rachises, apices acute or obtuse, margins serrate to biserrate, sinuses mostly 1–2(–4) mm deep; veins simple or 1–2-forked near the pinna bases, tips evident adaxially; indument abaxially of appressed tan clavate hairs 0.1 mm long; sori mostly 5–10 pairs per pinna, more numerous acroscopically than basiscopically; indusia 1.5–3 X 0.3–0.6 mm, margins entire; spores reniform.A
Forma de vida
Terrestre or epipetricA
Ejemplar revisado
Chis (Breedlove 22504, DS, MO, 69099, CAS). Gro (Velázquez 22, FCME). Oax (Hallberg 1493, NY, UC; Wendt et al. 5165, NY). Pue (Sánchez 73, US; Sánchez Mejorada R. 613, MEXU, US; ). Ver (Purpus 6198, UC, US; Ventura A. 11272, ASU, 14631, ARIZ, MO).A
Elevación
(500–)1000–2100(–2500) mA
Categoría IUCN
No incluidaC
Categoría NOM-059-SEMARNAT-2010
No incluidaD
Discusión taxonómica
See discussions under A. pteropus, A. barbaense, and A. potosinum for comparison with those species. Asplenium seileri, A. cruegeri, and A. pulchellum are additional relatives, albeit all smaller species. Asplenium miradorense is also sometimes confused with A. harpeodes, but the latter is epiphytic, at least in Mexico, while A. miradorense is strictly terrestrial. Asplenium miradorense lacks the long-attenuate pinna apices usually found in A. harpeodes. Elevationally, the two species appear nonoverlapping, or co-occurring only infrequently. Other closely related species are A. argentinum Hieron. (S. America), A. bangii Hieron. (Peru, Bolivia), A. boliviense Rosenst., A. brasilienseRaddi (S. America), A. claussenii Hieron. (Antilles, S. America), A. diplosceum Hieron. (Greater Antilles), A. otites Link, A. pulchellum Raddi, and A. raddianum Gaudich. Adams (in Davidse et al., 1995) included A. standleyi as a synonym of A. potosinum, but we believe it is a form of A. miradorense. This whole group, including A. miradorense, is badly in need of monographic revision, a fact about which all students of the group are in agreement.
Breedlove 31319 (DS, MEXU, NY), cited by Murakami and Moran (1993: 14) as A. laetum at NY (p.p.) and determined by Adams as A. cruegeri vel prox. aff., is neither species, but may be a hybrid between A. miradorense and A. abscissum. Most young sporangia are collapsed, and mature sporangia are seemingly without spores; rhizomes are erect (which precludes this specimen being A. laetum). Both putative parents are known from the area (Breedlove 31317, 31327, DS, A. miradorense; 31307, DS,A. abscissum), and the supposed hybrid is intermediate morphologically.
A
Breedlove 31319 (DS, MEXU, NY), cited by Murakami and Moran (1993: 14) as A. laetum at NY (p.p.) and determined by Adams as A. cruegeri vel prox. aff., is neither species, but may be a hybrid between A. miradorense and A. abscissum. Most young sporangia are collapsed, and mature sporangia are seemingly without spores; rhizomes are erect (which precludes this specimen being A. laetum). Both putative parents are known from the area (Breedlove 31317, 31327, DS, A. miradorense; 31307, DS,A. abscissum), and the supposed hybrid is intermediate morphologically.
A