Asplenium castaneum Schltdl. & Cham.

Primary tabs

Asplenium castaneum Schltdl. & Cham.

Descripción

Roots fibrous, not proliferous; rhizomes suberect; rhizome scales black, clathrate, not occluded, 4–6 X 0.3–0.6 mm, entire, tips long-attenuate; fronds clumped, numerous, mostly 12–35 cm long; stipes castaneous, lustrous, mostly 5–11(–17) cm X 0.7–2 mm, 1⁄4–1⁄2 of frond length, glabrous or with scattered hair-like scales (1–2 mm long) proximally, not alate; blades thickherbaceous to subcoriaceous, 1-pinnate, linear, 15–30 X 1–2 cm, apices pinnatifid, not proliferous; rachises reddish brown, with scattered filiform scales to ca. 1 mm and a few spreading septate hairs 0.2–0.4 mm, adaxial wings entire or rarely minutely papillate, 0.3 mm wide; pinnae nearly round or rhomboid, (10–)20–30 pairs, 6–10 X 5–6 mm, sessile, articulate, rachis color running out onto pinna stalks, the largest ones slightly auriculate acroscopically, margins entire to faintly crenulate; veins simple or 1-forked, tips evident adaxially; indument absent, blades essentially glabrous, or with sparse to scattered clavate hairs; sori 2–6 pairs per pinna, 2–4 on basiscopic sides, 2–5(–6) on acroscopic sides; indusia whitish to light tan, 1–3 X 0.8–1.2 mm, margins erose; spores globose to ovoid.A

Forma de vida

TerrestreA

Forma de crecimiento

Hierba

Nutrición

Autotrófica

Ejemplar revisado

Chis (Matuda 2353, K, MEXU, US). DF (Balls 4099, UC, US). Gro (Lorea 3980, 4205, 4360, FCME). Gto (Rzedowski 47913, IEB; 47913a, UC). Jal (Beaman 2372, UC, US). Méx (Beaman 2840, UC). Mich (Díaz B. 1009, ENCB, IEB, MEXU). Mor (Sánchez 310, MEXU). Oax (Mickel 4582, NY). Pue (Diggs 2060, ENCB, NY). Tlax (Acosta P. & Sánchez 2968, CIB). Ver (Beaman 2288, UC, US).
Unverified, Doubtful, or Mistaken Reports. Dgo (reported by Mickel & Beitel, 1988, but not verified; most likely, specimens so determined are A. sanchezii). NL (Hinton 17189, TEX, cited by Hinton & Hinton, 1995).
A

Distribución

Costa Rica presentB, Guatemala presentB, México (Country) native and not endemicB, Panama presentB, South America presentB

Elevación

(2450–)33004350 mA

Ecología y Hábitat

Often in rock crevices.A

Tipo de vegetación

Bosque de otro tipo

Tipo de vegetación (notas)

Usually above timberline, or in fir forests.A

Categoría IUCN

No incluidaC

Categoría NOM-059-SEMARNAT-2010

No incluidaD

Estatus del taxón

(A) Como definida actualmente, probablemente una entidad natural (monofilética)

Discusión taxonómica

Asplenium castaneum is distinguished from A. monanthes in having castaneous rachises and clathrate, non-occluded rhizome scales. Another related species is A. polyphyllum, but that has a subterminal proliferous bud along the rachis of each blade, the pinnae are more oblong, and the indusia subentire. Stolze (1986) and Tryon and Stolze (1993) treated A. castaneum as a variety of A. monanthes, but the two seem readily distinct and, in Mexico at least, seldom if ever grow together.
Occasional specimens of A. castaneum are very small, with as few as 8 pinna pairs (e.g., Ortega O. 538, XAL, from Veracruz). These specimens approach A. fibrillosum in size, but the indusia are subentire or erose and the rachises less scaly. Small specimens of A. castaneum also resemble A. sanchezii, which see for differences.
A

Bibliografía

A. Mickel, J. T. & Smith, A. R. 2004: The Pteridophytes of Mexico Vol. 88
B. Mickel, J. T. & Smith, A. R. 2004: The Pteridophytes of Mexico Vol. 88
C. IUCN 2022: The IUCN Red List of Threatened Species. Versión 2022-2
D. SEMARNAT 2019: MODIFICACIÓN del Anexo Normativo III, Lista de especies en riesgo de la Norma Oficial Mexicana NOM-059-SEMARNAT-2010: 101 pp. – https://www.dof.gob.mx/nota_detalle.php?codigo=5578808&fecha=14/11/2019#gsc.tab=0 [accessed 2023-05-04 06:16]