Cheilanthes longipila Baker
Descripción
Rhizomes compact, ascending, ca. 2 mm diam.; rhizome scales lanceolate, concolorous tan to orange-tan, 3–5 mm long, somewhat resinous, entire; fronds 12–30 cm long, clumped; stipes 1⁄3–2⁄5 the frond length, atropurpureous to black, terete, with abundant straight, patent, white to tan, gland-tipped hairs 0.5–2 (mostly ca. 1) mm long; blades narrowly deltate to lanceolate, tripinnatepinnatifid to quadripinnate; pinnae mostly opposite; segments mostly broadly adnate, nearly equilateral; adaxial surfaces densely clothed with nearly straight, somewhat appressed white hairs 1–3 mm long; abaxial surfaces densely clothed with flexuous white hairs; sori at vein tips, margin slightly or not curved, unmodified into false indusia; spores dark brown.A
Forma de vida
Epipétrica, TerrestreA
Distribución
México (país) EndémicoA
Elevación
(600 –)1400 – 1900 mA
Ecología y Hábitat
On rocky slopes.A
Tipo de vegetación
Bosque de encinoA
Categoría IUCN
No incluidaB
Categoría NOM-059-SEMARNAT-2010
No incluidaC
Discusión taxonómica
Key to the Mexican Varieties of Cheilanthes longipila
Adaxial segment hairs mostly 1–3 mm long, abaxially ca. 1 mm, flexuous; stipe hairs 0.5–2 mm; NL to Jal and Oaxaca. …......................................................................var. longipila.
Adaxial segment hairs ca. 0.5–1 mm long, abaxially ca. 0.5 mm, erect, straight; stipe hairs 0.2–0.5(–1) mm; Gro. .....................................................................................var. brevipila.
Cheilanthes longipila var. longipila Baker, Ann. Bot. (Oxford) 5(2): 211. 1891
Myriopteris longipila sub. longipila (Baker) Grusz & Windham, PhytoKeys 32: 58. 2013
Hemionitis longipila sub. longipila (Baker) Christenh., Global Fl. 4: 17. 2018
Differing from var. brevipila by its longer blade hairs and more widespread distribution; 2n=60 (NL).
Distribution. Dry, rocky slopes in oak forests; (600–)1450–1900 m. Mexico.
Selected Specimens Examined. Gro (Hinton et al. 14378, NY). Jal (Pringle 1863, NY; Palmer 78, NY; Pringle 5409, 2768, MEXU). Méx (Sánchez Mejorada 812, MEXU). NL (Correll & Johnston 19810, ASC, LL, NY). Oax (Mickel 1669, 3912, 6317, NY).
Unverified, Doubtful, or Mistaken Reports. Ags (reported by de la Cerda Lemus & Siqueiros Delgado, 1985, but not verified).
Var. longipila is distinct by the long rigid patent hairs and lanceolate to narrowly deltate blades. See C. maxoniana for comparison with that species.
Cheilanthes longipila var. brevipila Mickel, Mem. New York Bot. Gard. 88: 198 (-199; figs. 84N-Q, 87J-M). 2004
Myriopteris longipila subsp. brevipila (Mickel) Grusz & Windham, PhytoKeys 32: 58. 2013
A var. longipila pinnis alternatis et pilis laminarum brevioribus differt.
Rhizomes compact, ascending, 2–3 mm diam.; rhizome scales lanceolate, dull tan to orange, 3–4 mm long, entire; fronds to 30 cm long, clumped; stipes 1⁄3 the frond length, reddish brown, lustrous, terete, with patent glandular hairs mostly 0.2–0.5 mm long, some to 1 mm long; blades oblong to lanceolate, 2–3-pinnate-pinnatifid; pinnae alternate above the basal pair, nearly equilateral; rachises flattened, costae grooved; adaxial blade surfaces with scattered white ± appressed hairs ca. 1 mm long; abaxial blade surfaces with hairs ca. 0.5 mm, erect; sori on segment lobes, laminar margins curved, but not differentiated, with marginal hairs; spores dark brown.
Distribution. Tropical deciduous forest, oak woods, rocky slopes with volcanic soil; 1550–1600 m. Mexico.
Specimens Examined. Gro (Antonio 339, FCME, NY; Calzada 16172, NY).
This taxon is similar to C. longipila var. longipila, but has shorter, more variable hairs on all frond surfaces.A
Adaxial segment hairs mostly 1–3 mm long, abaxially ca. 1 mm, flexuous; stipe hairs 0.5–2 mm; NL to Jal and Oaxaca. …......................................................................var. longipila.
Adaxial segment hairs ca. 0.5–1 mm long, abaxially ca. 0.5 mm, erect, straight; stipe hairs 0.2–0.5(–1) mm; Gro. .....................................................................................var. brevipila.
Cheilanthes longipila var. longipila Baker, Ann. Bot. (Oxford) 5(2): 211. 1891
Myriopteris longipila sub. longipila (Baker) Grusz & Windham, PhytoKeys 32: 58. 2013
Hemionitis longipila sub. longipila (Baker) Christenh., Global Fl. 4: 17. 2018
Differing from var. brevipila by its longer blade hairs and more widespread distribution; 2n=60 (NL).
Distribution. Dry, rocky slopes in oak forests; (600–)1450–1900 m. Mexico.
Selected Specimens Examined. Gro (Hinton et al. 14378, NY). Jal (Pringle 1863, NY; Palmer 78, NY; Pringle 5409, 2768, MEXU). Méx (Sánchez Mejorada 812, MEXU). NL (Correll & Johnston 19810, ASC, LL, NY). Oax (Mickel 1669, 3912, 6317, NY).
Unverified, Doubtful, or Mistaken Reports. Ags (reported by de la Cerda Lemus & Siqueiros Delgado, 1985, but not verified).
Var. longipila is distinct by the long rigid patent hairs and lanceolate to narrowly deltate blades. See C. maxoniana for comparison with that species.
Cheilanthes longipila var. brevipila Mickel, Mem. New York Bot. Gard. 88: 198 (-199; figs. 84N-Q, 87J-M). 2004
Myriopteris longipila subsp. brevipila (Mickel) Grusz & Windham, PhytoKeys 32: 58. 2013
A var. longipila pinnis alternatis et pilis laminarum brevioribus differt.
Rhizomes compact, ascending, 2–3 mm diam.; rhizome scales lanceolate, dull tan to orange, 3–4 mm long, entire; fronds to 30 cm long, clumped; stipes 1⁄3 the frond length, reddish brown, lustrous, terete, with patent glandular hairs mostly 0.2–0.5 mm long, some to 1 mm long; blades oblong to lanceolate, 2–3-pinnate-pinnatifid; pinnae alternate above the basal pair, nearly equilateral; rachises flattened, costae grooved; adaxial blade surfaces with scattered white ± appressed hairs ca. 1 mm long; abaxial blade surfaces with hairs ca. 0.5 mm, erect; sori on segment lobes, laminar margins curved, but not differentiated, with marginal hairs; spores dark brown.
Distribution. Tropical deciduous forest, oak woods, rocky slopes with volcanic soil; 1550–1600 m. Mexico.
Specimens Examined. Gro (Antonio 339, FCME, NY; Calzada 16172, NY).
This taxon is similar to C. longipila var. longipila, but has shorter, more variable hairs on all frond surfaces.A