Cheilanthes pyramidalis Fée
Descripción
Rhizomes compact, horizontal, 2–3 mm diam.; rhizome scales linear-subulate, bicolorous, dark brown to black with narrow light brown entire margins, lustrous, ca. 3 mm long; fronds 15–45 cm long, clumped; stipes 1⁄2–2⁄3 the frond length, castaneous, lustrous, grooved, glabrous; blades 3–4-pinnate-pinnatifid, ovate to deltate, rarely narrowly oblong; segments narrowly obcuneate to linear, obtuse, often strongly drooping; adaxial surfaces with 0.1–0.3 mm long white papillae at segment apices and on false indusia; abaxial surfaces glabrous; laminar margins recurved, strongly modified into 0.3–0.5 mm wide false indusia covering most of abaxial laminar surfaces, margins long-fimbriate (hairs 0.3 mm), generally long-decurrent along pinnule axes; spores grayish brown; n=2n=87, apogamous.A
Forma de vida
Rupícola, TerrestreB
Ejemplar revisado
Ags (McVaugh & Koelz 212, MICH). BCS (Carter & Ferris 3360A, MEXU). Chih (Pringle 1442, NY). Chis (Breedlove 22395, NY). Col (Reko 4847, US). DF (Rusby 365, NY). Dgo (Pennell 18342, NY). Gro (Hinton 9650, NY). Gto (Dugas s.n., in 1880, NY). Hgo (Rose 9205, NY). Jal (Mexia 1513, NY). Méx (Mickel 697, NY). Mich (Hinton 13421, NY). Mor (Clark 7286, NY). Nay (Norris & Taranto 14636, MICH). Oax (Mickel 3891, NY). Pue (Riba et al. 324, MEXU). Qro (Díaz B. 4972a, IEB). SLP (Wendt &Chiang 12273d, NY). Sin (Breedlove 16748, NY). Son (Gentry et al. 19321, NY). Tlax (Vibrans 400D, MEXU). Ver (Dorantes et al. 05113, NY). Zac (McVaugh 17760, NY).A
Elevación
850 – 3000 mA
Ecología y Hábitat
Grassy slopes and rocks in dry woods to moist woods.A
Tipo de vegetación
Bosque de pino-encinoA
Categoría IUCN
No incluidaC
Categoría NOM-059-SEMARNAT-2010
No incluidaD
Discusión taxonómica
Cheilanthes pyramidalis represents one of the most difficult complexes in Mexican pteridophytes. It is one of the most widespread ferns in Mexico, occurring in 23 states and the Distrito Federal, and displaying a wide range of variation. Typically, the species has fimbriate, decurrent indusia with spicules on the indusial surfaces and along the laminar-indusial interfaces or even onto the adaxial blade surfaces. The segments may be narrowly oblong or, on occasion, linear and drooping. Some specimens lack spicules, or the indusia may not be decurrent. The blades are often lanceolate but may be broadly deltate.
The name C. hirsuta Link has been generally applied to this species in recent years, but the isotype at PH has the longdecurrent indusia of C. marginata. As the specific epithet suggests, it is unique in having hairs on the segments, not only at the indusial margin but especially along the indusial-laminar interfaces, with some of the hairs irregularly broadened (scale-like). Such hairs are not found in specimens of C. pyramidalis, but are occasional in C. marginata. Thus, we consider C. hirsuta a synonym of C. marginata. Cheilanthes marginata is clearly allied and often confused with C. pyramidalis. Typically, it too has fimbriate, long-decurrent indusia, but it generally lacks spicules, the marginal fimbriae are longer, and the indusia are broader (0.5 mm) and longerdecurrent, not just to the subtending axis, but to the next segment. Blades of C. marginata are usually deltate (occasionally lanceolate) with less elongate segments, and the species generally occurs at higher elevations than does C. pyramidalis.A
The name C. hirsuta Link has been generally applied to this species in recent years, but the isotype at PH has the longdecurrent indusia of C. marginata. As the specific epithet suggests, it is unique in having hairs on the segments, not only at the indusial margin but especially along the indusial-laminar interfaces, with some of the hairs irregularly broadened (scale-like). Such hairs are not found in specimens of C. pyramidalis, but are occasional in C. marginata. Thus, we consider C. hirsuta a synonym of C. marginata. Cheilanthes marginata is clearly allied and often confused with C. pyramidalis. Typically, it too has fimbriate, long-decurrent indusia, but it generally lacks spicules, the marginal fimbriae are longer, and the indusia are broader (0.5 mm) and longerdecurrent, not just to the subtending axis, but to the next segment. Blades of C. marginata are usually deltate (occasionally lanceolate) with less elongate segments, and the species generally occurs at higher elevations than does C. pyramidalis.A