Cheilanthes complanata A.R.Sm.
Descripción
Rhizomes compact, horizontal to ascending, 2–3 mm diam.; rhizome scales linear-lanceolate, bicolorous, black with narrow brown margin, lustrous, 3–4 mm long, entire; fronds 30–42 cm long, clumped; stipes 1⁄2–2⁄3 the frond length, castaneous, terete or slightly grooved distally, glabrous; blades deltate, attenuate at apices, tripinnate to tripinnate-pinnatifid; pinnae ca. 15 pairs, equilateral except the basiscopically exaggerated basal pair; segments narrowly obovate to narrowly elliptic; adaxial surfaces pebbly, but without papillae; abaxial surfaces glabrous except for scattered, minute, orange, 1-celled hairs; laminar margins recurved, somewhat differentiated into 0.3 mm wide, smooth false indusia, entire, not or only slightly decurrent along next axis; spores gray-brown.A
Forma de vida
Epipétrica, TerrestreA
Distribución
México (país) Nativo y no endémicoA
Elevación
1960 – 3000 mA
Tipo de vegetación
Bosque de neblina/mesófiloA
Categoría IUCN
No incluidaB
Categoría NOM-059-SEMARNAT-2010
No incluidaC
Discusión taxonómica
Cheilanthes complanata is distinct by its non-decurrent, entire indusia, castaneous stipes and rachises, bicolorous rhizome scales, and conspicuous swollen vein endings (generally visible abaxially). It is close to C. harrisii Maxon of Jamaica in its nondecurrent, entire, barely modified false indusia and deltateattenuate blades, but C. harrisii differs in its rhizome scales being somewhat flexuous with kinky hair-tips, frond axes darker (atropurpureous vs. castaneous), and blades more dissected (to quadripinnate vs. tripinnate to tripinnate-pinnatifid).
Key to the Mexican Varieties of Cheilanthes complanata
Indusia continuous, unlobed; Chis. .....................var. complanata.
Indusia discontinuous, deeply lobed; Gro. ............var. interrupta.
Cheilanthes complanata var. complanata A.R.Sm., Amer. Fern J. 70: 19, figs. 1980
Var. complanata differs from var. interrupta in having continuous (vs. discontinuous) indusia.
Distribution. Terrestrial or epipetric in evergreen cloud forests, rocky slopes in pine woods; 2750–3000 m. Mexico.
Specimens Examined. Chis (Breedlove 40335, DS, 55870, CAS, ENCB, NY).
Unverified, Doubtful, or Mistaken Reports. Ver (reported by Mickel & Beitel, 1988, but not verified).
At least some of the specimens cited by Yatskievych and Moran (in Davidse et al., 1995) are here treated as C. decurrens, which see.
Cheilanthes complanata var. interrupta Mickel, Mem. New York Bot. Gard. 88: 189 (fig. 77J-L). 2004
A C. complanata var. complanata indusiis lobatis vel discretis distincta. (Alluding to the indusia being interrupted along the segment margins, rather than continuous as in the type variety.)
Rhizomes compact, ascending, 2–3 mm diam.; rhizome scales linear-lanceolate, 3–4 mm long, bicolorous with a black central stripe with brown margins, lustrous, entire; fronds to 42 cm long, clumped; stipes 2⁄3 the frond length, dark reddish brown, lustrous, grooved, glabrous; blades deltate, attenuate at apices, tripinnate at base; pinnae ca. 15 pairs, equilateral except basal pair which is exaggerated basiscopically, segments oblong, obtuse to acute; adaxial surfaces glabrous; abaxial surfaces glabrous except for sparse orangish 1-celled hairs; laminar margins recurved, somewhat differentiated as false indusia, 0.3–0.5 mm wide, lobed, discontinuous at segment apices, not at all decurrent; spores gray-brown.
Distribution. Slopes in laurel forests; 1960 m. Known only from the type collection.
Var. interrupta resembles var. complanata in its narrowly deltate-attenuate blades, castaneous blade axes, and bicolorous rhizome scales, but differs in the indusia being lobed to discrete,
rather than entire.
A
Key to the Mexican Varieties of Cheilanthes complanata
Indusia continuous, unlobed; Chis. .....................var. complanata.
Indusia discontinuous, deeply lobed; Gro. ............var. interrupta.
Cheilanthes complanata var. complanata A.R.Sm., Amer. Fern J. 70: 19, figs. 1980
Var. complanata differs from var. interrupta in having continuous (vs. discontinuous) indusia.
Distribution. Terrestrial or epipetric in evergreen cloud forests, rocky slopes in pine woods; 2750–3000 m. Mexico.
Specimens Examined. Chis (Breedlove 40335, DS, 55870, CAS, ENCB, NY).
Unverified, Doubtful, or Mistaken Reports. Ver (reported by Mickel & Beitel, 1988, but not verified).
At least some of the specimens cited by Yatskievych and Moran (in Davidse et al., 1995) are here treated as C. decurrens, which see.
Cheilanthes complanata var. interrupta Mickel, Mem. New York Bot. Gard. 88: 189 (fig. 77J-L). 2004
A C. complanata var. complanata indusiis lobatis vel discretis distincta. (Alluding to the indusia being interrupted along the segment margins, rather than continuous as in the type variety.)
Rhizomes compact, ascending, 2–3 mm diam.; rhizome scales linear-lanceolate, 3–4 mm long, bicolorous with a black central stripe with brown margins, lustrous, entire; fronds to 42 cm long, clumped; stipes 2⁄3 the frond length, dark reddish brown, lustrous, grooved, glabrous; blades deltate, attenuate at apices, tripinnate at base; pinnae ca. 15 pairs, equilateral except basal pair which is exaggerated basiscopically, segments oblong, obtuse to acute; adaxial surfaces glabrous; abaxial surfaces glabrous except for sparse orangish 1-celled hairs; laminar margins recurved, somewhat differentiated as false indusia, 0.3–0.5 mm wide, lobed, discontinuous at segment apices, not at all decurrent; spores gray-brown.
Distribution. Slopes in laurel forests; 1960 m. Known only from the type collection.
Var. interrupta resembles var. complanata in its narrowly deltate-attenuate blades, castaneous blade axes, and bicolorous rhizome scales, but differs in the indusia being lobed to discrete,
rather than entire.
A