Cheilanthes peninsularis Maxon
Descripción
Rhizomes short-creeping, horizontal, 1 mm diam.; rhizome scales linear-lanceolate, concolorous orange-brown, weakly bicolorous with age, weakly and irregularly toothed to entire; fronds to 26(–30) cm, clumped; stipes 1⁄2 the frond length, castaneous to purplish brown, grooved, lustrous, with scattered, filiform, tortuous to straight, orange-brown scales, 1–3 mm, plus many reduced, often contorted and hair-tipped scales/hairs, 0.1–0.3 mm; blades deltate to lanceolate, tripinnate-pinnatifid; rachises and rachillae with scales like those of stipes (but lesser axes with deltate-attenuate and tortuous scales); pinnae 5–11, equilateral except basal pair basiscopically exaggerated; adaxial surfaces green, glabrous; abaxial surfaces glabrous except for scales on dark axes; sori along segment tips only, margins curved, but not differentiated; spores tan.A
Forma de vida
Epipétrica, TerrestreA
Distribución
México (país) EndémicoA
Elevación
0 – 1350 mA
Ecología y Hábitat
On rocky slopes among shrubs and in open woods.A
Categoría IUCN
No incluidaB
Categoría NOM-059-SEMARNAT-2010
No incluidaC
Discusión taxonómica
Key to the Mexican Varieties of Cheilanthes peninsularis
Rachilla scales flexuous, denticulate; blades deltate, 3-pinnate-pinnatifid to 4-pinnate; pinna pairs ca. 6; BCS. ..............................................................................................................var. peninsularis.
Rachilla scales nearly straight, subentire; blades lanceolate, 2–3-pinnate; pinna pairs 7–11; Rev. .....................................................................................................................................var. insularis.
Cheilanthes peninsularis var. peninsularis Maxon, Contr. U.S. Natl. Herb. 10: 496. 1908
Myriopteris peninsularis subsp. peninsularis (Weath.) Grusz & Windham, PhytoKeys 32: 60. 2013
Differing from var. insularis in having flexuous, denticulate rachillae scales, deltate, tripinnate-pinnatifid to quadripinnate blades, and ca. 6 pinna pairs.
Distribution. Rocky open woods, often on shaded north slopes; 575–1350 m. Mexico.
Selected Specimens Examined. BCS (Brandegee 660, GH, NY, UC; Carter & Moran 5507, UC; Jones 24559, CAS, NY; Moran 7093, DS; Porter 253, DS; Pray 1787, 1793, LAM; Wiggins 5465, CAS, 14763, 15377, DS; Wiggins et al. 495, DS, UC).
Two specimens, Pray 1760 and 1761, LAM, seem to be C. peninsularis var. peninsularis, but are labelled as coming from San Luis Potosí, where this species is not known to occur. We suspect they were mislabelled since other specimens of C. peninsularis var. peninsularis were collected by Pray in Baja California.
Cheilanthes peninsularis var. peninsularis is distinct from C. pringlei in the former having slender purplish brown stipes with fewer (scattered) narrower scales (vs. stout reddish brown stipes with copious scales), fronds narrower ovate (vs. short, triangular or deltate-ovate), pinnae spaced (vs. close and imbricate), and rachises and rachillae with sparse yellow-brown scales (vs. with numerous broad whitish scales extending thickly even to distal parts of the pinnules, commonly obscuring the abaxial surfaces).
Cheilanthes peninsularis var. insularis Weath., Amer. Fern J. 21: 25. 1931
Myriopteris peninsularis subsp. insularis (Weath.) Grusz & Windham, PhytoKeys 32: 60. 2013
Differing from the type variety in having nearly straight, subentire rachilla scales, lanceolate, 2–3-pinnate blades, and 7–11 pinna pairs.
Distribution. Rock crevices and volcanic talus, dry scrub; 0–1025 m. Mexico.
Selected Specimens Examined. BCS (Brandegee s.n., 17 Feb 1889, NY, is somewhat intermediate). Rev (Cruz Cisneros 1603, ENCB; Dawson 13272, UC; Elmore C8, DS; Felger 15794, NY, SD, UC; Howell 8393, CAS, GH, NY; Lindsay 2836, SD; Mason 14593, CAS, MEXU, UC; Moran 5718, SD, 5928, DS, SD; Villareal 15, ENCB).A
Rachilla scales flexuous, denticulate; blades deltate, 3-pinnate-pinnatifid to 4-pinnate; pinna pairs ca. 6; BCS. ..............................................................................................................var. peninsularis.
Rachilla scales nearly straight, subentire; blades lanceolate, 2–3-pinnate; pinna pairs 7–11; Rev. .....................................................................................................................................var. insularis.
Cheilanthes peninsularis var. peninsularis Maxon, Contr. U.S. Natl. Herb. 10: 496. 1908
Myriopteris peninsularis subsp. peninsularis (Weath.) Grusz & Windham, PhytoKeys 32: 60. 2013
Differing from var. insularis in having flexuous, denticulate rachillae scales, deltate, tripinnate-pinnatifid to quadripinnate blades, and ca. 6 pinna pairs.
Distribution. Rocky open woods, often on shaded north slopes; 575–1350 m. Mexico.
Selected Specimens Examined. BCS (Brandegee 660, GH, NY, UC; Carter & Moran 5507, UC; Jones 24559, CAS, NY; Moran 7093, DS; Porter 253, DS; Pray 1787, 1793, LAM; Wiggins 5465, CAS, 14763, 15377, DS; Wiggins et al. 495, DS, UC).
Two specimens, Pray 1760 and 1761, LAM, seem to be C. peninsularis var. peninsularis, but are labelled as coming from San Luis Potosí, where this species is not known to occur. We suspect they were mislabelled since other specimens of C. peninsularis var. peninsularis were collected by Pray in Baja California.
Cheilanthes peninsularis var. peninsularis is distinct from C. pringlei in the former having slender purplish brown stipes with fewer (scattered) narrower scales (vs. stout reddish brown stipes with copious scales), fronds narrower ovate (vs. short, triangular or deltate-ovate), pinnae spaced (vs. close and imbricate), and rachises and rachillae with sparse yellow-brown scales (vs. with numerous broad whitish scales extending thickly even to distal parts of the pinnules, commonly obscuring the abaxial surfaces).
Cheilanthes peninsularis var. insularis Weath., Amer. Fern J. 21: 25. 1931
Myriopteris peninsularis subsp. insularis (Weath.) Grusz & Windham, PhytoKeys 32: 60. 2013
Differing from the type variety in having nearly straight, subentire rachilla scales, lanceolate, 2–3-pinnate blades, and 7–11 pinna pairs.
Distribution. Rock crevices and volcanic talus, dry scrub; 0–1025 m. Mexico.
Selected Specimens Examined. BCS (Brandegee s.n., 17 Feb 1889, NY, is somewhat intermediate). Rev (Cruz Cisneros 1603, ENCB; Dawson 13272, UC; Elmore C8, DS; Felger 15794, NY, SD, UC; Howell 8393, CAS, GH, NY; Lindsay 2836, SD; Mason 14593, CAS, MEXU, UC; Moran 5718, SD, 5928, DS, SD; Villareal 15, ENCB).A