Amauropelta resinifera (Desv.) Pic.Serm.
Descripción
Rhizomes erect; fronds (15–)25–100(–130) cm long; stipes stramineous to tan, usually mucilaginous when young, 2–10(–25) cm X 1–3(–6) mm diam., mostly less than 10(–25) cm long; blades to 110 X 4–25(–40) cm, with to 12 pairs of gradually reduced proximal pinnae, lowermost pinnae hastate to auriculate; pinnae to 12(–20) X (0.4–)0.7–1.5(–2) cm (excluding basal auricles), ascending in the distal third of blades, incised to within 1 mm of costae; aerophores short, peg-like at pinna bases, or seemingly absent; segments usually very oblique to the costae, often falcate, deltate to lanceolate, to ca. 2.5 mm wide; veins (3–)5–10 pairs per segment; indument abaxially of usually short hairs mostly less than 0.3(–0.5) mm (occasionally glabrous) on rachises, costae, and costules, tissue between veins often shorthairy and usually with numerous golden to reddish, sessile, hemispherical, resinous glands; sori medial to supramedial, with persistent tan usually glandular indusia; sporangia glabrous; 2n=58 (Ver, var. resinifera), 116 (Jam).A
Forma de vida
TerrestreA
Elevación
100 – 2300 mA
Tipo de vegetación
Bosque de pino-encino, Bosque de otro tipoA
Categoría IUCN
No incluidaB
Categoría NOM-059-SEMARNAT-2010
No incluidaC
Discusión taxonómica
(In relation to the synonym Thelypteris resinifera)
Key to the Mexican Varieties of Thelypteris resinifera
Segments slightly oblique, ± straight; blades narrowly elliptic; western Mexico (Chihuahua to Jalisco)……………………………...…………...…………...…………...…………...…….var. proxima
Segments strongly oblique and falcate; blades broadly elliptic; e. and s. Mexico, w. Mexico n. to Jalisco...............................................................…………...…………..........var. resinifera
Thelypteris resinifera var. resinifera (Desv.) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 63. 1953
Nephrodium panamense C.Presl, Reliq. Haenk. 1(1): 35. 1825
Polypodium litigiosum Liebm., Kongel. Danske Vidensk.-Selsk. Skr. V(1): 205 (seors. 53). 1849
Aspidium exsudans E.Fourn., Mexic. Pl. 1: 93. 1872
Aspidium exsudans var. majus E.Fourn., Mexic. Pl. 1: 93. 1872
Aspidium ghiesbreghtii E.Fourn., Mexic. Pl. 1: 94. 1872
Dryopteris panamensis C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. ser. 7, 4: 292. 1907
Differing from var. proxima in the more broadly elliptic, generally larger blades and the more strongly oblique and falcate segments.
Distribution. In woods and clearings, especially along trails and near streams, also in seepage areas and roadside ditches; 100–1600(–2300) m. USA (Fla); Mexico; Guat, Bel, Hond, Salv, Nic, CR, Pan; Gr Ant; Col, Ec.
Selected Specimens Examined. Chis (Breedlove 31263, DS). Col (Lott 2988 et al., UC). Gro (Tenorio L. 460, CAS, ENCB, MEXU). Jal (McVaugh 20461, IEB, MEXU). Méx (Hinton 7323, GH, K, LL, MEXU, NY, US). Mich (Torres C. 363, ARIZ, ENCB, MEXU). Mor (Salas M. 33, IEB). Oax (Mexia 9203, ARIZ, BH, CAS, GH, LL, MICH, NY, U, UC, US). Pue (Sánchez 641, MEXU). Qro (Rzedowski 42839, IEB, XAL). Tab (Linden 1526, K). Ver (Muñoz 16130, UC).
Unverified, Doubtful, or Mistaken Reports. Ags (De la Cerda & García 762, HUAA, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified).
This species is closely related to T. struthiopteroides and T. Balbisii (which see for further discussion), and a few specimens are difficult to place. Thelypteris resinifera can be distinguished by the combination of numerous, rather closely placed, gradually reduced proximal pinnae, sessile glands on the tissue between the veins abaxially, inequilateral pinnae with oblique, often falcate segments, and persistent indusia. There is considerable variation in pubescence: some specimens are nearly glabrous abaxially (e.g., Medina A. & Vázquez B. 371, XAL, from Veracruz), but most specimens have rather dense hairs on costae, veins, and tissue between veins. Indusia vary from glabrous to hairy, but most have at least a few hairs.
Thelypteris resinifera var. proxima (C.Chr.) A.R.Sm., Mem. New York Bot. Gard. 88: 634. 2004
Dryopteris panamensis var. proxima C.Chr., Smithsonian Misc. Collect. 52: 377. 1909
Differing from var. resinifera by the characters mentioned in the key.
Distribution. Terrestrial in moist pine-oak forests, especially along streams and seepage areas, or in streambeds; 1000–1900 m. Mexico.
Selected Specimens Examined. Chih (Knobloch 579, US). Jal (Rose & Painter 7320, US; Sada 16163, UC). Mor (Ross 326, BM, M). Nay (Alava & Cook 1623, UC; Tenorio L. 16793, MEXU). Sin (Breedlove 35694, CAS; González O. 5070, MEXU, US).
Mexia 608 (CAS, F, GH, UC), from Nayarit, and Salazar 287 (US), from Sinaloa, are atypical in the dense septate hairs on the rachises and costae abaxially. They do not differ from other specimens of this variety in frond shape, size, and dissection.
Additional study is needed to determine whether the type of var. proxima, from Morelos, is really convarietal with the other specimens of this distinctive variety.A
Key to the Mexican Varieties of Thelypteris resinifera
Segments slightly oblique, ± straight; blades narrowly elliptic; western Mexico (Chihuahua to Jalisco)……………………………...…………...…………...…………...…………...…….var. proxima
Segments strongly oblique and falcate; blades broadly elliptic; e. and s. Mexico, w. Mexico n. to Jalisco...............................................................…………...…………..........var. resinifera
Thelypteris resinifera var. resinifera (Desv.) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 63. 1953
Nephrodium panamense C.Presl, Reliq. Haenk. 1(1): 35. 1825
Polypodium litigiosum Liebm., Kongel. Danske Vidensk.-Selsk. Skr. V(1): 205 (seors. 53). 1849
Aspidium exsudans E.Fourn., Mexic. Pl. 1: 93. 1872
Aspidium exsudans var. majus E.Fourn., Mexic. Pl. 1: 93. 1872
Aspidium ghiesbreghtii E.Fourn., Mexic. Pl. 1: 94. 1872
Dryopteris panamensis C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. ser. 7, 4: 292. 1907
Differing from var. proxima in the more broadly elliptic, generally larger blades and the more strongly oblique and falcate segments.
Distribution. In woods and clearings, especially along trails and near streams, also in seepage areas and roadside ditches; 100–1600(–2300) m. USA (Fla); Mexico; Guat, Bel, Hond, Salv, Nic, CR, Pan; Gr Ant; Col, Ec.
Selected Specimens Examined. Chis (Breedlove 31263, DS). Col (Lott 2988 et al., UC). Gro (Tenorio L. 460, CAS, ENCB, MEXU). Jal (McVaugh 20461, IEB, MEXU). Méx (Hinton 7323, GH, K, LL, MEXU, NY, US). Mich (Torres C. 363, ARIZ, ENCB, MEXU). Mor (Salas M. 33, IEB). Oax (Mexia 9203, ARIZ, BH, CAS, GH, LL, MICH, NY, U, UC, US). Pue (Sánchez 641, MEXU). Qro (Rzedowski 42839, IEB, XAL). Tab (Linden 1526, K). Ver (Muñoz 16130, UC).
Unverified, Doubtful, or Mistaken Reports. Ags (De la Cerda & García 762, HUAA, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified).
This species is closely related to T. struthiopteroides and T. Balbisii (which see for further discussion), and a few specimens are difficult to place. Thelypteris resinifera can be distinguished by the combination of numerous, rather closely placed, gradually reduced proximal pinnae, sessile glands on the tissue between the veins abaxially, inequilateral pinnae with oblique, often falcate segments, and persistent indusia. There is considerable variation in pubescence: some specimens are nearly glabrous abaxially (e.g., Medina A. & Vázquez B. 371, XAL, from Veracruz), but most specimens have rather dense hairs on costae, veins, and tissue between veins. Indusia vary from glabrous to hairy, but most have at least a few hairs.
Thelypteris resinifera var. proxima (C.Chr.) A.R.Sm., Mem. New York Bot. Gard. 88: 634. 2004
Dryopteris panamensis var. proxima C.Chr., Smithsonian Misc. Collect. 52: 377. 1909
Differing from var. resinifera by the characters mentioned in the key.
Distribution. Terrestrial in moist pine-oak forests, especially along streams and seepage areas, or in streambeds; 1000–1900 m. Mexico.
Selected Specimens Examined. Chih (Knobloch 579, US). Jal (Rose & Painter 7320, US; Sada 16163, UC). Mor (Ross 326, BM, M). Nay (Alava & Cook 1623, UC; Tenorio L. 16793, MEXU). Sin (Breedlove 35694, CAS; González O. 5070, MEXU, US).
Mexia 608 (CAS, F, GH, UC), from Nayarit, and Salazar 287 (US), from Sinaloa, are atypical in the dense septate hairs on the rachises and costae abaxially. They do not differ from other specimens of this variety in frond shape, size, and dissection.
Additional study is needed to determine whether the type of var. proxima, from Morelos, is really convarietal with the other specimens of this distinctive variety.A