Anemia tomentosa (Sav.) Sw.
Categoría IUCN
No incluidaA
Categoría NOM-059-SEMARNAT-2010
No incluidaB
Discussion
Anemia tomentosa var. mexicana (C.Presl) Mickel, Iowa State J. Sci. 36: 427. 1962
Anemia distans Fée, Mém. Foug. 9 Cat. Foug. Mex.: 41. 1857
Anemia fulva var. mexicana C.Presl, Suppl. Tent. Pterid. 84. 1845
Rhizomes horizontal, compact, short-creeping, ca. 7 mm diam.; rhizome hairs orange; fronds erect, 9–39 cm tall; stipes 1⁄2–2⁄3 the frond length, 0.9–2.2 mm diam., stramineous to light brown, hirsute; blades deltate-ovate, bipinnate-pinnatifid, 4.6–10.7 cm wide, chartaceous to subcoriaceous; pinnae 6–13 pairs, opposite to subopposite, pinnules narrowly adnate, ovate, segment lobes acute; blade surfaces hirsute; veins free; fertile pinnae approximate to or remote from the sterile pinnae, about equal to the sterile blades in height; spores striate, ridges smooth, close; 2n=152 (Jal), 2n=228 (Oax).
Distribution. Infrequent on open to slightly shaded grassy slopes, often grazed, generally disturbed; 390–1850 m, down to 60 m in Isthmus of Tehuantepec. Mexico; Hisp; Col, Ven. Three other varieties are known from southern South America.
Selected Specimens Examined. Chih (Palmer 91, NY). Dgo (Ortega 4294, US). Gto (Ventura V. & López P. 9888, IEB). Jal (Pringle 1834, NY). Méx (Kimnach & Moran 139, UC). Nay (Jones 23473, UC). Oax (Mickel 761, NY). Pue (Purpus 4045, UC). Sin (Breedlove 18050, NY). SLP (Schaffner 26, NY). Son (Gentry et al. 19290, NY). Tam (Bartlett 10866, MICH). Ver (Purpus 8460, NY). Zac (Anderson 12631, NY).
Unverified, Doubtful, or Mistaken Reports. Ags (González-Adame 612, HUAA, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified). BCN (reported by Knobloch & Correll, 1962, and Wiggins, 1980, as Anemia anthriscifolia).
The two cytotypes of A. tomentosa in Mexico apparently are allopatric and can be distinguished by spore size. The hexaploid, which occurs in Oaxaca, has spores 76–94 (avg. ca. 81) µm in diameter, whereas material from Jalisco (tetraploid) and Sonora has spores 69–79 (avg. ca. 74) µm in diameter.
Anemia tomentosa is distinguished in Mexico by its thin texture and bipinnate-pinnatifid blades. In Mexico it most closely resembles A. karwinskyana, which is distinct in its catadromous architecture and has segments that are more obtuse and rounded rather than toothed.
C
Anemia distans Fée, Mém. Foug. 9 Cat. Foug. Mex.: 41. 1857
Anemia fulva var. mexicana C.Presl, Suppl. Tent. Pterid. 84. 1845
Rhizomes horizontal, compact, short-creeping, ca. 7 mm diam.; rhizome hairs orange; fronds erect, 9–39 cm tall; stipes 1⁄2–2⁄3 the frond length, 0.9–2.2 mm diam., stramineous to light brown, hirsute; blades deltate-ovate, bipinnate-pinnatifid, 4.6–10.7 cm wide, chartaceous to subcoriaceous; pinnae 6–13 pairs, opposite to subopposite, pinnules narrowly adnate, ovate, segment lobes acute; blade surfaces hirsute; veins free; fertile pinnae approximate to or remote from the sterile pinnae, about equal to the sterile blades in height; spores striate, ridges smooth, close; 2n=152 (Jal), 2n=228 (Oax).
Distribution. Infrequent on open to slightly shaded grassy slopes, often grazed, generally disturbed; 390–1850 m, down to 60 m in Isthmus of Tehuantepec. Mexico; Hisp; Col, Ven. Three other varieties are known from southern South America.
Selected Specimens Examined. Chih (Palmer 91, NY). Dgo (Ortega 4294, US). Gto (Ventura V. & López P. 9888, IEB). Jal (Pringle 1834, NY). Méx (Kimnach & Moran 139, UC). Nay (Jones 23473, UC). Oax (Mickel 761, NY). Pue (Purpus 4045, UC). Sin (Breedlove 18050, NY). SLP (Schaffner 26, NY). Son (Gentry et al. 19290, NY). Tam (Bartlett 10866, MICH). Ver (Purpus 8460, NY). Zac (Anderson 12631, NY).
Unverified, Doubtful, or Mistaken Reports. Ags (González-Adame 612, HUAA, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified). BCN (reported by Knobloch & Correll, 1962, and Wiggins, 1980, as Anemia anthriscifolia).
The two cytotypes of A. tomentosa in Mexico apparently are allopatric and can be distinguished by spore size. The hexaploid, which occurs in Oaxaca, has spores 76–94 (avg. ca. 81) µm in diameter, whereas material from Jalisco (tetraploid) and Sonora has spores 69–79 (avg. ca. 74) µm in diameter.
Anemia tomentosa is distinguished in Mexico by its thin texture and bipinnate-pinnatifid blades. In Mexico it most closely resembles A. karwinskyana, which is distinct in its catadromous architecture and has segments that are more obtuse and rounded rather than toothed.
C