Adiantum andicola Liebm.
Descripción
Rhizomes compact to long-creeping, 2–5 mm diam.; rhizome scales usually black, or castaneous only at rhizome tips, appressed, 1–1.5 X 0.1–0.3 mm, margins entire; fronds clumped to often well spaced, (23–)35–70 cm long, arching; stipes atropurpureous, lustrous, 10–40 cm X 1–3 mm, 1⁄2–2⁄3 the frond length, glabrous except at bases; blades broadly ovate to deltate, 3–5-pinnate proximally, 15–50 X 15–60 cm, tapering and less divided towards apices; rachises atropurpureous, glabrous, sometimes glaucous; pinnae 4–8 compound pairs, alternate, lowermost stalked 2–7 cm; pinnulets obovate to rhombic or flabellate, narrowly (to broadly) cuneate at bases, tiny (4 X 4 mm) to large (22 X 25 mm), distal margins incised-lobate, often articulate; veins free, forking, ending in teeth (when teeth are present); indument absent on both sides of blades, abaxially often glaucous and with or without a dense covering of minute papillae; idioblasts absent; sori 2–8 per pinnulet, confined to distal margins; indusia 1–2 mm long, reniform, round-reniform, or lunate, glabrous.A
Forma de vida
TerrestreA
Ejemplar revisado
Chih (Jenkins et al. s.n., 27 July 1988, MO). Chis (Breedlove 32455, DS, NY). DF (Purpus 1584, NY, UC, US). Dgo (Palmer 176, K, NY, US). Gro (Cowan 4948, CAS, NY, TEX, UC). Gto (Díaz Barriga 3054, ENCB, IEB). Hgo (Gimate 753, ENCB, NY). Jal (McVaugh 26137, NY). Méx (Pringle 15638, CAS, UC, US). Mich (Hitchcock & Stanford 7142, DS, NY). Mor (Copeland herb. 113, MEXU, NY, UC). Nay (Boutin & Brandt 2646, CAS). NL (Meyer & Rogers 3057, MO). Oax (Mickel 6150, ENCB, NY). Pue (Orcutt 4025, DS, US). Qro (Rubio 2117, MEXU). SLP (Rzedowski 27710, IEB). Sin (Vega et al. 3014, ENCB, FCME). Tam (Hinton 24893, TEX). Tlax (Chimal et al. 28, IEB). Ver (García & Delgado 950, CHAPA, MO, NY, XAL).
Unverified, Doubtful, or Mistaken Reports. Ags (De la Cerda & García 1350, HUAA, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified). Tab (reported by Magaña, 1992, probably based on a misidentification).
A
Unverified, Doubtful, or Mistaken Reports. Ags (De la Cerda & García 1350, HUAA, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified). Tab (reported by Magaña, 1992, probably based on a misidentification).
A
Elevación
1000 – 3150 mA
Tipo de vegetación
Bosque de encino, Bosque de pino-encino, Bosque de neblina/mesófilo, Selva alta, Bosque de otro tipoA
Categoría IUCN
No incluidaB
Categoría NOM-059-SEMARNAT-2010
No incluidaC
Discusión taxonómica
Adiantum andicola is most likely a complex comprising several species and is in need of monographic study. The papillate nature of the surface of pinnulets in some variants has been overlooked, and is most evident on specimens from Oaxaca and Guerrero. There is considerable variation in this species in pinnulet size, shape, blade dissection, glaucousness, and rhizome form (compact to long-creeping), but some correlation exists between pinnulet size and papilla size. The variants lumped together here have in common dark, usually black, rhizome scales (not found in other species with articulate pinnulets). Maxon and Morton’s unpublished A. hintonii (represented by King & Soderstrom 5043, NY; Rzedowski 45333, IEB; both from Michoacán), with larger, broader pinnulets, is one of the more distinct variants; it is most common in western and central Mexico but extends to Panama. Other variants have more deeply incised pinnulets. Breedlove 42784 (DS), from Chiapas, also has more flabellate pinnulets and wider indusial sinuses, and Balls B4703 (UC), from Veracruz, has more lunate sori that lack pronounced sinuses, reminiscent of sori in A. braunii. Another extreme, with numerous, small, narrow pinnulets, and large laminar papillae (longer than wide), appears to be most common in Oaxaca, Veracruz, Chiapas, and Guerrero, but other small-pinnuleted specimens from these states and elsewhere in Mexico lack papillae. Specimens from Hidalgo and Tamaulipas are atypical in having more rounded, less incised pinnulets; identifications of several of these (e.g., Hernández Magaña 6092, MEXU; Hinton 24893, TEX) are problematic.A