Notholaena standleyi Maxon

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Notholaena standleyi Maxon

Descripción

Rhizomes compact, horizontal, 1–1.5 mm diam.; rhizome scales lanceolate, 2–3 mm long, bicolorous with broad dark brown to black, sclerotic, lustrous central streak and pale entire margins; fronds 9–23 cm long, clumped; stipes ca. 3⁄4 the frond length, castaneous, terete, lustrous, glabrous except for a few broadly lanceolate scales at bases (2 X 1 mm); blades broadly pentagonal, 2.5–7 cm wide, pinnatifid above basal pinnae, with the suprabasal pinnae usually shortened, basal pinnae pinnatifid with one basiscopically exaggerated pinnatifid basal pinnule; adaxial surfaces glabrous; abaxial surfaces densely pale yellow-to deep yellow-farinose; laminar margins slightly curved; sporangia 32- or 16-spored; spores tan to black; 2n=60 (Oax, USA).A

Forma de crecimiento

Hierba

Forma de vida

Epipétrica, TerrestreB

Nutrición

Autotrófica

Ejemplar revisado

BCN (Moran 18714, ENCB). BCS (Carter 2049, MEXU, US). Chih (Pringle 467, NY). Coah (Palmer 1379, NY). Dgo (Purpus 4632, US). Hgo (Rose 9107, US). NL (Rodríguez L. & Sauceda M. 412, ANSM). Oax (Mickel 4490, NY). Pue (Purpus 4027, NY). Sin (Pitelka 286B, UC). SLP (Parry 992, NY). Son (Lumholtz Exped. 361, NY). Tam (Bartlett 10708, US).A

Distribución

México (país) Nativo y no endémicoA, Norteamérica al N de México PresenteA

Elevación

380 – 2200(– 3100) mA

Ecología y Hábitat

Calcareous or noncalcareous cliffs, igneous rocks, sandstone, rock crevices, among large boulders and cliffs, limestone outcrops.A

Tipo de vegetación

Matorral rosetófiloB

Categoría IUCN

No incluidaC

Categoría NOM-059-SEMARNAT-2010

No incluidaD

Estatus del taxón

(A) Como definida actualmente, probablemente una entidad natural (monofilética)

Discusión taxonómica

The farina varies in color geographically. In Sonora, Chihuahua, and part of Coahuila the farina is very pale yellow, to nearly white in some Chihuahuan specimens. In part of Coahuila and in San Luis Potosí, the farina is deep yellow, and in Puebla and Oaxaca pale yellow.
Windham (in FNA Ed. Comm., 1993) reported that the low number of spores per sporangium in N. standleyi is apparently not associated with apogamy since all plants analyzed were sexual diploids. Siegler and Wollenweber (1983) identified three chemotypes in this species; these correlate with substrate specificity, range, and variations in farina color.
A

Bibliografía

A. Mickel, J. T. & Smith, A. R. 2004: The Pteridophytes of Mexico Vol. 88
B. Velázquez Montes, E. 2019: Familia Pteridaceae. En: Rzedowski, J. & Hernández Ledesma, P. (Eds.). Fl. Bajío Regiones Adyacentes Fasc. 210: 1-249
C. IUCN 2022: The IUCN Red List of Threatened Species. Versión 2022-2
D. SEMARNAT 2019: MODIFICACIÓN del Anexo Normativo III, Lista de especies en riesgo de la Norma Oficial Mexicana NOM-059-SEMARNAT-2010: 101 pp. – https://www.dof.gob.mx/nota_detalle.php?codigo=5578808&fecha=14/11/2019#gsc.tab=0 [accessed 2023-05-04 06:16]