Cystopteris fragilis (L.) Bernh.
Descripción
Rhizomes short-creeping, not protruding past youngest leaves; rhizome scales brown, lanceolate, lustrous, membranous, entire, mixed with fine hairs; fronds to 47 cm long, clumped; stipes 1⁄3–1⁄2 the frond length, stramineous, often reddish brown and with scales at bases, glabrous or with scattered glandular hairs; blades (6–)9–20 X (2.5–)4–8 cm, bipinnate to tripinnate-pinnatifid, ovate-lanceolate with proximal 1–2 pairs of pinnae slightly shortened; rachises glabrous to pubescent with glandular hairs; pinnae 7–15 pairs, 1–4 cm long, thin, but more than 2 cells thick; segments oblong, 1.5–4 mm wide; veins ending in small emargination or in teeth; indusia glabrous with subentire to lacerate apices; 2n=168 (Chis).A
Forma de vida
Epipétrica, TerrestreA
Ejemplar revisado
BCS (Carter et al. 2336, DS, UC). Chih (Correll & Gentry 23207, MO, UC). Chis (Breedlove 15111, DS). Coah (Wynd & Mueller 521, MO). DF (Bourgeau 251, MO). Dgo (Tenorio L. et al. 4159, MEXU). Gro (Mexia 9068, MO, UC). Gto (Ventura & López 2758, IEB). Hgo (Arreola 5, FCME). Jal (Díaz-Luna 6027, GUADA, UC). Méx (Fanti 1037, ENCB, FCME, MO). Mich (Díaz-Luna 9292, GUADA, UC). Mor (Hinton 17521 bis, ENCB, MEXU). Nay (McVaugh 18888, IEB, MEXU, MICH, MO). NL (Hinton et al. 17304, MO). Oax (Mickel 5529, NY, UC). Pue (Orcutt 3983, MO). Qro (Zamudio 6894, IEB). Sin (Breedlove 44959, CAS). SLP (Palmer 176, MO, UC). Tam (Lof 705, UC). Ver (Ventura A. 17697, MEXU).
Unverified, Doubtful, or Mistaken Reports. Ags (Siqueiros 4346, HUAA, and three other collections, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified).
A
Unverified, Doubtful, or Mistaken Reports. Ags (Siqueiros 4346, HUAA, and three other collections, cited by Siqueiros-Delgado & González-Adame, 2004, but not verified).
A
Distribución
Asia PresenteA, Caribe PresenteA, Centroamérica: Costa Rica PresenteA; El Salvador PresenteA; Guatemala PresenteA; Honduras PresenteA; Nicaragua PresenteA; Panamá PresenteA, Europa PresenteA, México (país) Nativo y no endémicoA, Norteamérica al N de México PresenteA, Sudamérica PresenteA, África PresenteA
Elevación
(600 –)1050 – 3400(– 4050) mA
Ecología y Hábitat
Often on or among rocks, on mossy ledges.A
Tipo de vegetación
Selva baja, Bosque de encino, Bosque de neblina/mesófilo, Bosque de pino-encinoA
Categoría IUCN
Preocupación menor (LC)B
Categoría NOM-059-SEMARNAT-2010
No incluidaC
Discusión taxonómica
Until the Mexican members of the Cystopteris fragilis complex have been studied and characterized to the degree that taxa north of Mexico have been (chromosomes, isozymes, etc.), a conservative treatment is followed, one that combines C. fragilis s.str. with C. diaphana. The North American taxa include diploids, allopolyploids, and various sterile hybrids, with species often having limited ranges. Cystopteris reevesiana Lellinger of the southwestern United States probably occurs in northern Mexico but is not readily distinguishable from Mexican C. fragilis and is not treated here. It is supposed to be distinguished from C. fragilis by blades bipinnate-pinnatifid to tripinnate (vs. pinnatepinnatifid to bipinnate-pinnatifid) and rhizomes usually longcreeping (vs. short-creeping). These characters do not seem to be correlated in Mexican specimens, and more work is needed to circumscribe the species.
Cystopteris fragilis most closely resembles a small Dryopteris, but the hood-like indusia of the former readily distinguish it.
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Cystopteris fragilis most closely resembles a small Dryopteris, but the hood-like indusia of the former readily distinguish it.
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