Sticherus bifidus (Willd.) Ching

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Sticherus bifidus (Willd.) Ching

Descripción

Rhizomes long-creeping, with castaneous to dark brown ciliate scales 2–3 X 0.2–0.3 mm; stipes 2–3 mm diam.; main axes dormant or often continuing beyond first fork, scaly to glabrescent; dormant buds with tan to whitish, lax, long-ciliate scales 1.5–2 X 0.2–0.4 mm; pinnae 1–2 times pseudodichotomously forking; axes below ultimate forks pectinate on acroscopic side or lacking segments altogether; penultimate segments 3–8 cm wide, the costae bearing tan to whitish ciliate scales 1.5–2 X 0.2–0.3 mm; ultimate segments chartaceous to subcoriaceous, linear, to ca. 4 cm X 2–3 mm at mid-segment, plane to usually revolute, abaxially densely tomentose with arachnoid hairs; sori inframedial to medial, with 3–4(–5) sporangia per sorus; 2n=68 (Chis, Jam, PR).A

Forma de crecimiento

Hierba

Forma de vida

TerrestreA

Nutrición

Autotrófica

Ejemplar revisado

Chis (Breedlove 34600, DS, ENCB). Gro (Cowan 4921, ENCB, NY, UC). Jal (McVaugh 21405, IEB, MEXU, MICH, NY). Méx (Tejero-Díez 3171, IZTA). Oax (Mickel 1026, ENCB, MEXU, NY, 5911, ENCB, MEXU, NY, UC). Pue (Marquez R. et al. 756, ENCB, MEXU, XAL; Ventura A. 374, ENCB, NY). Tab (Cowan 3171, MEXU, NY). Ver (Pringle 6130, BR, ENCB, K, MEXU, MO, NY, UC, US).A

Distribución

Caribe PresenteA, Centroamérica: Costa Rica PresenteA; El Salvador PresenteA; Guatemala PresenteA; Honduras PresenteA; Nicaragua PresenteA; Panamá PresenteA, México (país) Nativo y no endémicoA, Sudamérica PresenteA

Elevación

150 – 2100 mA

Ecología y Hábitat

Common on roadside banks, clearings and forest margins, forming dense masses and thickets in lower montane and montane forests.A

Tipo de vegetación

Selva alta, Bosque de neblina/mesófilo, Bosque de encinoA

Categoría IUCN

No incluidaB

Categoría NOM-059-SEMARNAT-2010

No incluidaC

Estatus del taxón

(A) Como definida actualmente, probablemente una entidad natural (monofilética)

Discusión taxonómica

This is the commonest species of Sticherus in Mexico and is readily identified by its abaxially dense indument, which often hides the sori. It is most easily confused with S. brevipubis, which has similar dense arachnoid indument on the blades abaxially, but in that species (as well as in S. palmatus) the secondary axes (immediately proximal to the ultimate forks) are often pectinate on both sides of the axis, whereas in S. bifidus the secondary axes are seldom wholly pectinate; however, both species may be pectinate only on the acroscopic side. The dormant bud scales in S. brevipubis are dark brown, rigid, and mostly short-ciliate, and the costal scales on abaxial surfaces are nearly absent and less than 1 mm long.
Gonzales (2003) has adopted the name Sticherus ferrugineus (Desv.) J. Gonzales, comb. ined., for this species, based on Mertensia ferruginea Desv., Ges. Naturf. Freunde Berlin Mag. Neusten Entdeck. Gesamten Naturk. 5: 307 (1811). Type. French Guiana, collector unknown (holotype: P, herb. A. N. Desvaux). Gonzales gave as distribution for S. ferrugineus: Mexico to Panama, Greater and Lesser Antilles, Venezuela, Colombia to Bolivia, and southeastern Brazil. According to Gonzales, S. bifidus s. str. co-occurs with S. ferrugineus in the southern part of this range (Panama, Cuba, Colombia to Bolivia, Venezuela, Trinidad, Guyana, French Guiana, and southeastern Brazil) and differs primarily by having narrower bud scales with truncate bases, lacking aphlebiae and proximal internal segments on the basal 1–2 cm of each axis, having a matted-whitish indument, and having laxer, more appressed axis scales.
A

Bibliografía

A. Mickel, J. T. & Smith, A. R. 2004: The Pteridophytes of Mexico Vol. 88
B. IUCN 2022: The IUCN Red List of Threatened Species. Versión 2022-2
C. SEMARNAT 2019: MODIFICACIÓN del Anexo Normativo III, Lista de especies en riesgo de la Norma Oficial Mexicana NOM-059-SEMARNAT-2010: 101 pp. – https://www.dof.gob.mx/nota_detalle.php?codigo=5578808&fecha=14/11/2019#gsc.tab=0 [accessed 2023-05-04 06:16]