Ceradenia L.E. Bishop
Descripción
Rhizomes short-creeping or erect, dorsiventral or radially symmetrical; rhizome scales orangish to castaneous or atropurpureous, non-clathrate, usually setulose or with marginal glands (rarely, scales absent: C. jungermannioides); fronds monomorphic, determinate (ours) or indeterminate; stipes often articulate, bearing setae and, abaxially, usually branched hairs; blades rarely simple, commonly pinnatifid, pinnatisect, or perpinnate; costae not readily visible abaxially; veins usually obscure, pinnate, simple or often 1-forked from the costae; hydathodes absent, vein tips not visible adaxially; indument of reddish brown to atropurpureous, stiff setae often 1–3 mm long on stipes, rachises, costae, and sometimes laminae, also of shorter branched or unbranched hairs, these often septate and sometimes bearing whitish globose or oblong glands, usually 2–5 glands per hair; sori round or slightly oblong, superficial or slightly impressed; indusia absent; soral paraphyses present, in the form of hairs bearing whitish to occasionally tan or yellowish wax-like globose or pyriform glands that function to protect the developing sporangia (and obscure them in young sori); sporangia glabrous; spores tetrahedral-globose; x=37.A
Forma de vida
Epífita at times epipetric or terrestrial at high elevation.A
Categoría IUCN
No incluidaD
Categoría NOM-059-SEMARNAT-2010
No incluidaE
Discusión taxonómica
Ceradenia is widespread in the wet Neotropics and comprises about 55 species; a single species (C. jungermannioides) extends to the Azores, and another (C. pruinosa) is probably conspecific with the African species Ctenopteris punctata F. Ballard. The genus is most closely related to Enterosora (which see) and probably also to Zygophlebia, with about eight, mostly neotropical species. Recent molecular work on Grammitidaceae by Ranker et al. (2004) indicates that Ceradenia is very likely monophyletic.
Soral paraphyses, whitish glands, and anhydathodous blades distinguish Ceradenia from other Mexican genera of Grammitidaceae. The glands are usually globose or pear-shaped and are borne terminally and laterally on short uniseriate hairs; the glands may completely hide the young sporangia and can be mistaken for spores when sporangia mature. Specimen-drying treatments, especially if an alcohol-based preservative is used before drying, may obscure or apparently destroy the glands. Some species in subg. Ceradenia (e.g., C. pruinosa) also have branched gland-bearing hairs on the laminar tissue, on the abaxial surface or on both surfaces. Often there are three (2–5) globose or oblong glands per hair.
In Mexico and usually wherever they grow, species of Ceradenia are infrequent or rare, often very local, and usually found only in relatively undisturbed cloud forests with constant high humidity. There are relatively few collections of all four Mexican species.
A
Soral paraphyses, whitish glands, and anhydathodous blades distinguish Ceradenia from other Mexican genera of Grammitidaceae. The glands are usually globose or pear-shaped and are borne terminally and laterally on short uniseriate hairs; the glands may completely hide the young sporangia and can be mistaken for spores when sporangia mature. Specimen-drying treatments, especially if an alcohol-based preservative is used before drying, may obscure or apparently destroy the glands. Some species in subg. Ceradenia (e.g., C. pruinosa) also have branched gland-bearing hairs on the laminar tissue, on the abaxial surface or on both surfaces. Often there are three (2–5) globose or oblong glands per hair.
In Mexico and usually wherever they grow, species of Ceradenia are infrequent or rare, often very local, and usually found only in relatively undisturbed cloud forests with constant high humidity. There are relatively few collections of all four Mexican species.
A