Athyrium bourgaei E. Fourn.
Description
Similar to A. arcuatum except: rhizome scales pale brown, 9–11 X 1–1.5 mm; stipes 7–47 cm long, 1⁄3–1⁄2 the frond length; blades 18–55 X 7–18 cm, linear to linear-lanceolate, basal pinnae often deflexed; basal pinnae (2.5–)6–9.5 X (1.5–)2.5–3 cm, basal basiscopic pinnule short, 0.6–0.8 cm long, largest basiscopic pinnule 1.6–2.5 cm long, basiscopic and acroscopic pinnules roughly equal in length, costae abaxially with sparse to scattered pale brown, hair-like scales 1–1.5 X 0.1–0.3 mm, pinna margins slightly recurved, serrate; wings along costae and segment midribs conspicuous adaxially, with short teeth 0.1 mm at vein branchings; indusia 0.5–0.8 mm long, entire or sparingly fimbriate.A
Forma de vida
TerrestreA
Specimen
BCS (Brandegee s.n., 3 Oct 1899, UC). Chih (Correll & Gentry 23079, NY, UC). Chis (Breedlove 21998, NY). DF (Lyonnet 2682, MEXU). Dgo (Benítez P. 2282a, CHAPA, MEXU). Gto (Rzedowski 44805, IEB, MEXU). Hgo (Moore 5378, UC). Jal (Equipo 4-15475, GUADA, UC). Méx (Rzedowski 30903, NY). Mich (Bartholomew 2862, CAS, NY). Mor (Sandoval 46, MEXU). Nay (Breedlove 44488, CAS, MEXU). NL (Rodríguez L. 135, ANSM). Oax (Mickel 7433, NY, UC). Pue (Flores O. 122, ENCB). Qro (Díaz-Barriga & Carranza 7285, NY). Son (Van Devender 97-850, ARIZ). Ver (Johnson s.n., 18 Sep 1906, NY).A
Altitude
1850 - 3600 mA
Habitat & Ecology
In moist woods and along streams.A
Tipo de vegetación
Bosque de pino, Bosque de encino, Bosque de neblina/mesófiloA
Categoría IUCN
No incluidaB
Categoría NOM-059-SEMARNAT-2010
No incluidaC
Discussion
This taxon was referred by Smith (1981) and Stolze (1981) to the north temperate species Athyrium filix-femina (L.) Roth, sensu lato, which is in need of modern taxonomic monographic work. We can find no distinct break between A. bourgeaui of Mexico and A. filix-femina vars. californicum and cyclosorum of the western United States, but until the relationships of the North American and European species are understood, we prefer to use the earliest Mexican name for this taxon. A similar case is made for using the earliest Mexican name (Athyrium arcuatum) for the taxon referred to by Smith (1981) as A. dombei Desv., a species described from Peru. Athyrium bourgeaui has lanceolate fronds, narrower than those of A. arcuatum, basal pinnae smaller and more deflexed, longer and broader rhizome scales, acroscopic and basiscopic pinnules roughly equal in size, and sparse to scattered, hair-like scales along the costae. Athyrium arcuatum has deltate-lanceolate fronds broader than those of A. bourgeaui, larger ascending to spreading basal pinnae, short and narrow rhizome scales, basiscopic pinnules longer than acroscopic pinnules, and glabrous costae. Athyrium barnebyanum, with scattered black, hair-like scales on the abaxial laminar surfaces, costae, and rachises, and dark brown to black scales 2.5–3 mm wide on rhizomes and stipe bases, resembles A. arcuatum in blade size and shape, and is here included in the latter species.
A
A